My publications

Below is a list of my publications in order of release from oldest to newest; they include peer-reviewed articles, survey reports and articles related to various aspects of herpetological research in magazines or non-peer reviewed media.

Published Research and Observations

1. Allain, S. J. R. & Goodman, M. J. (2014). Evaluation of great crested newt and smooth newt populations at Bar Hill Crematorium. Cambridgeshire Bird Report 2013, 87, 156-161.
2. Allain, S. J. R. & Goodman, M. J. (2015). Cambridgeshire Amphibian Survey Report 2014. Nature in Cambridgeshire, 57, 101-108.
3. Stark, T., Viernum, S., Allain, S., Spiller, M. & Reeves, V. (2015). Crowdfunding for Chytrid 2.0 (Batrachochytrium salamandrivorans) in Belgium. Froglog, 116, 20.
4. Allain, S. J. R. & Goodman, M. J. (2016). Lamination as a method of preserving reptile sloughs. Herpetological Bulletin, 138, 29-30.
5. Allain, S. J. R. & Smith, L. T. (2016). Newt mortalities on an urban cycle path. Herpetological Bulletin, 138, 27-28.
6. Allain, S. J. R. & Goodman, M. J. (2017). Cambridgeshire Amphibian Survey Report 2015. Nature in Cambridgeshire, 59, 35-44.
7. Allain, S. J. R., Smith, L. T. & Miller, G. J. (2017). Introduced non-native amphibians and reptiles in Cambridgeshire 2010-2016. Nature in Cambridgeshire, 59, 44-50.
8. Allain, S., Stark, T. & Viernum, S. (2017). An Update on Crowdfunding for Chytrid 2.0. FrogLog, 118, 29.
9. Allain, S. J. R. & Goodman, M. J. (2017). A case of xanthochromism in the common frog (Rana temporaria). Herpetological Bulletin, 139, 39-40.
10. Allain, S. J. R. & Goodman, M. J. (2017). Using call playbacks to investigate a population of non-native midwife toads Alytes obstetricans (Laurenti, 1768) in Cambridge, UK. Herpetological Bulletin, 140, 28-30.
11. Allain, S. J. R. & Smith, L. T. (2017). New records of paedomorphic smooth newts (Lissotriton vulgaris) at a site in Cambridgeshire, U.K. Herpetological Bulletin, 141, 40.
12. Allain, S. J. R. & Goodman, M. J. (2017). Monitoring a Non-native Amphibian in Cambridge, UK. FrogLog, 119, 36-37.
13. Allain, S. J. R. (2017). The use of roadside gabions as refugia by reptiles in Kota Kinabalu, Sabah, Malaysia. Captive & Field Herpetology, 1(1), 7-9.
14. Allain, S. J. R. & Goodman, M. J. (2017). Absence of chytrid fungus (Batrachochytrium dendrobatidis) in an introduced population of the common midwife toad (Alytes obstetricans) in Cambridge, UK. Herpetological Bulletin, 142, 40-41.
15. Allain, S. J. R. (2018). Comments on the Swabbing of Anuran Tadpoles for the Detection of Infectious Diseases. FrogLog, 120, 40.
16. Allain, S. J. R. & Goodman, M. J. (2018). The absence of the amphibian chytrid fungi in the common midwife toad (Alytes obstetricans) from an introduced population in Cambridge, UK. Herpetology Notes, 11, 451-454.
17. Allain, S. J. R. & Goodman, M. J. (2018). Cambridge Amphibian Survey Report 2016. Nature in Cambridgeshire, 60, 31-36.
18. Allain, S. J. R. & Goodman, M. J. (2018). New records of Palmate Newts (Lissotriton helveticus) in Cambridge. Nature in Cambridgeshire, 60, 36-37.
19. Allain, S. J. R. & Goodman, M. J. (2018). Digital measuring spoons, another instrument in the field herpetologist’s tool-box? Herpetological Bulletin, 144, 26-27.
20. Cunningham, O. K. E. & Allain, S. J. R. (2018). A Description of the Successful Hatching of the Fijian Banded Iguana (Brachylophus bulabula) and Observations on Vermiculite Moisture. Captive & Field Herpetology, 2(1), 12-15.
21. Allain, S. J. R. (2018). In the shadow of mammals: the use of camera traps to investigate the basking behaviour of Agama mwanzae in the Serengeti National Park, Tanzania. Herpetological Bulletin, 145, 19-21.
22. Allain, S. J. R. & Bateman, T. C. B. (2018). Ixodid Ticks on Oertzen’s Rock Lizard (Anatololacerta oertzeni) on Ikaria, Greece, with Notes on the Island’s Reptiles. IRCF Reptiles & Amphibians, 25(3), 176-179.
23. Allain, S. J. R. & Goodman, M. J. (2018). The Defense Mechanism of Kuhl’s Stream Frog (Limnonectes cf. kuhlii). IRCF Reptiles & Amphibians, 25(3), 180-181.
24. Allain, S. J. R. (2019). Now you see me, now you don't. Frontiers in Ecology & the Environment, 17(1), 31.
25. Saucedo, B., Garner, T. W. J., Kruithof, N., Allain, S. J. R., Goodman, M. J., Cranfield, R. J., Sergeant, C., Vergara, D. A., Kik, M. J. L., Forzán, M. J., van Beurden, S. J. & Gröne, A. (2019). Common midwife toad ranaviruses replicate first in the oral cavity of smooth newts (Lissotriton vulgaris) and show distinct strain-associated pathogenicity. Scientific Reports, 9, 4453.
26. Allain, S. J. R., Gandola, R. & Wilkinson, J. W. (2019). One or multiple origins of midwife toads in the UK? FrogLog, 121, 23.
27. Allain, S. J. R. (2019). Comments on the collection of buccal swabs from amphibians. FrogLog, 121, 29.
28. Allain, S. J. R., Goodman, M. J. & Jopling, A. D. (2019). Notes on the successful mitigation-driven translocation of slow worms (Anguis fragilis) at Wandlebury Country Park. Nature in Cambridgeshire, 61, 67-69.
29. Allain, S. J. R. & Goodman, M. J. (2019). New records of midwife toads (Alytes obstetricans) in Cambridgeshire. Nature in Cambridgeshire, 61, 69-70.
30. Allain, S. J. R. & Goodman, M. J. (2019). Cambridge Amphibian Survey Report 2017. Nature in Cambridgeshire, 61, 70-75.
31. Cunningham, O. K. E. & Allain, S. J. R.. (2019). First record of intraspecific oophagy in the common leaf-tailed gecko Uroplatus fimbriatus (SCHNEIDER, 1797). Captive & Field Herpetology, 3(1), 19-21.
32. Allain, S. J. R. (2019). Mining Flickr: a method for expanding the known distribution of invasive species. Herpetological Bulletin, 148, 11-14.
33. Allain, S. J. R. & Smith, L. T. (2019). A suspected case of infection by Amphibiocystidium sp. at a site in Cambridgeshire, UK. SAURIA, 41(3), 58–60.
34. Allain, S. J. R. & Mattea, R. G. (2019). An observation of extreme necrosis in the Italian wall lizard (Podarcis sicula). IRCF Reptiles & Amphibians, 26(2), 102.
35. Bray, I. H. & Allain, S. J. R. (2019). Are squash boxes necessary? The use of dorsal photographs to determine snout-vent lengths of slow worms (Anguis fragilis). IRCF Reptiles & Amphibians, 26(2), 165-166.
36. Allain, S. J. R. & Duffus, A. L. J. (2019). Emerging Infectious Disease Threats to European Herpetofauna. Herpetological Journal. 29(4), 189-206.
37. Oberhofer, S. J. & Allain, S. J. R. (2019). Autumn activity in an introduced population of Podarcis muralis ssp. in Liechtenstein. L@CERTIDAE, 2019(7), 54–57.
38. Smales, L. R., Allain, S. J. R., Wilkinson, J. W. & Harris, E. (2020). A new species of Pseudoacanthocephalus (Acanthocephala: Echinorhynchidae) from the guttural toad, Sclerophrys gutturalis (Bufonidae), introduced into Mauritius, with comments on the implications of the introductions of toads and their parasites into the United Kingdom. Journal of Helminthology, 94, e119.
39. Allain, S. J. R. & Goodman, M. J. (2020). Notes on the Herpetofauna of the Singapore Botanic Gardens. IRCF Reptiles & Amphibians, 26(3), 216–218.
40. Allain, S. J. R. & Goodman, M. J. (2020). An Incidence of Road Mortality in the Asian Water Monitor (Varanus salvator) in Kota Kinabalu, Sabah, Malaysia. IRCF Reptiles & Amphibians, 26(3), 219–220.
41. Grassby-Lewis, R. J. & Allain, S. J. R. (2020). Observation of Arboreal Behavior in the Malayan Pitviper (Calloselasma rhodostoma). IRCF Reptiles & Amphibians, 27(1), 79–80.
42. Mattea, R. G. & Allain, S. J. R. (2020). An Observation of Nocturnal Activity in a Central-Italian Asp (Vipera aspis francisciredi). IRCF Reptiles & Amphibians, 27(1), 87–88.
43. Clemens, D. J. & Allain, S. J. R. (2020). New records of Aesculapian snakes (Zamenis longissimus) in South Wales, UK. Herpetological Bulletin, 152, 30-31.
44. Clemens, D. J. & Allain, S. J. R. (2020). First evidence of Fogo skink (Chioninia fogoensis) introduction to Madeira. Herpetological Bulletin, 152, 40-41.
45. Harnkness, H. & Allain, S. J. R. (2020). A case of leucism in the slow worm (Anguis fragilis). Herpetological Bulletin, 152, 43.
46. Thomas, O., Allain, S. J. R. & Sagar, P. (2020). Predation and ingestion of a viviparous lizard (Zootoca vivipara) by the common wall lizard (Podarcis muralis) in Dorset, UK. Herpetological Bulletin, 152, 44.
47. West, P. & Allain, S. J. R. (2020). A case of erythrism in the common frog (Rana temporaria). IRCF Reptiles & Amphibians, 27(2), 331–332.
48. Shin, Y., Yikweon, J., Allain, S. J. R. & Borzée, A. (2020). Catalogue of herpetological specimens of the Ewha Womans University Natural History Museum (EWNHM), Republic of Korea. ZooKeys, 965, 103–139.
49. Allain, S. J. R.. (2020) Snakes in the grass: the misidentification of adders. Nature in Cambridgeshire, 62, 51-54.
50. Allain, S. J. R. & Bray, I. H. (2020). The ongoing monitoring of slow worms (Anguis fragilis) at Wandlebury Country Park. Nature in Cambridgeshire, 62, 54-56.
51. Allain, S. J. R. & Garcia, G. (2020). Promoting research, education and conservation: 9th World Congress of Herpetology. Herpetological Bulletin, 153, 1-2.
52. Allain, S. J. R., Gandola, R., Tighe, A. & Wilkinson, J. W. An investigation into the provenance of Bedford’s midwife toads. Bedfordshire Naturalist, In Press.
53. Baker, J. M. R. & Allain, S. J. R. (2020). Photographic Identification of Individual Barred Grass Snakes Natrix helvetica. In Press.
54. Allain, S. J. R. & Goodman, M. J. Investigating the presence of the amphibian chytrid fungus in a non-native species: An update. FrogLog, In Press.
55. Allain, S. J. R., Goodman, M. J., Gandola, R. & Wilkinson, J. W. One or multiple origins of midwife toads in the UK? An update. FrogLog, In Press.

My citations

Academic Research Projects

1. Allain, S. J. R. (2015). 'The use of bottle traps at a colony of great crested newts (Triturus cristatus) at a site in Cambridgeshire to investigate aquatic microhabitat preferences', Bachelor's Dissertation, Anglia Ruskin University.
2. Allain, S. J. R. (2017). 'An expanded geographical range of the amphibian chytrid fungus (Batrachochytrium dendrobatidis) in Madagascar', Master's Thesis, Imperial College London.
3. Allain, S. J. R. (2018). 'Investigating the conservation status and changes in species temporal distributions of Asian amphibians using museum specimens', Master's Thesis, Imperial College London.

Magazine Articles

Coming soon!

Features

1. Allain, S. J. R. (2018). The making of an amphibian conservation biologist. BES Bulletin, 49(1), 64-65.
2. Price, J. (2018). Working for nature: Steven Allain - Amphibian and reptile surveys, Cambridgeshire. BBC Wildlife, 36(11), 115.

Please note: PDFs have been provided in favour of links, where possible.

ResearchGate Logo Google Scholar Logo ORCID Logo Mendeley Logo SCOPUS Logo

You can find out more about my current research by following me on the respective networks by clicking on the images above, visiting my blog or by visiting the Research page. If you'd like to contact me in regards to a potential collaboration then please do this via the Contact page.